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Proconsul (mammal): Difference between revisions - Wikipedia

Proconsul (mammal): Difference between revisions

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'''''Proconsul''''' is an [[extinct]] [[genus]] of [[primate]]s that existed from 21 to 1417 million years ago during the [[Miocene]] epoch. [[Fossil]] remains are present in Eastern [[Eastern Africa]], including [[Kenya]] and [[Uganda]]. Four species have been classified to date: [[Proconsul africanus|''P. africanus'']], ''P. gitongai'', ''[[Proconsul major|P. major]]'' and ''P. meswae''. The four species differ mainly in body size. Environmental reconstructions for the Early Miocene ''Proconsul'' sites are still tentative and range from forested environments to more open, arid grasslands.
 
The [[gibbon]] and [[great apes]], including humans, are held in evolutionary biology to share a common ancestral lineage, which may have included ''Proconsul''. Its name, meaning "before Consul" (Consul being a certain chimpanzee that, at the time of the genus's discovery, was on display in [[London]]), implies that it is ancestral to the [[Common chimpanzee|chimpanzee]]. It might also be ancestral to the rest of the [[ape]]s.
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The genus had a mixture of [[Old World monkey]] and [[ape]] characteristics, so its placement in the ape superfamily [[Hominoidea]] is tentative, with some scientists placing ''Proconsul'' outside it, before the split of the [[apes]] and [[Old World monkeys]].
 
''Proconsul's'' monkey-like features include [[Quadrupedalism#Pronograde posture|pronograde posturesposture]], indicated by a long flexible back, curved metacarpals, and an above-branch [[arboreal]] quadrupedal positional repertoire. The primary feature linking ''Proconsul'' with extant apes is its lack of a tail; other "ape-like" features include its enhanced grasping capabilities, stabilized elbow joint and facial structure. ''Proconsul'' could not [[Suspensory behavior|hang effortlessly]] from tree branches like gibbons and other nonhuman apes do today.
 
==Discovery and classification==
[[File:Consul-the-chimp.jpg|thumb|150px|Consul at the Belle Vue zoo, Manchester, c.1894]]
The first specimen, a partial jaw discovered in 1909 by a gold prospector at [[Koru, Kenya|Koru]], near [[Kisumu]] in western Kenya, was also the oldest fossil hominoid known until recently, and the first fossil mammal ever found in sub-Saharan Africa. The name, ''Proconsul'', was devised by Arthur Hopwood in 1933 and means "before Consul"; the name of a famous captive chimpchimpanzee in London.<ref name="Morell">{{Harvnb|Morell|1996| p=130}}</ref> At the time Consul was being used as a circus name for performing chimpanzees. The [[Folies Bergère]] of 1903 in [[Paris]] had a popular performing [[Common chimpanzee|chimpanzee]] named Consul, and so did the [[Belle Vue ZooZoological Gardens]] in [[Manchester]], England, in 1894. On the latter's death in that year [[Ben Brierley]] wrote a commemorative poem wondering where the "[[Transitional fossil|Missing Link]]" between chimpanzees and men was.<ref>{{Harvnb|Walker|Shipman|2005}}</ref>
 
[[File:Proconsul skull at AMNH.jpg|thumb|Skull of ''Proconsul africanus'' at the [[American Museum of Natural History]]]]
Hopwood in 1931 had discovered the fossils of three individuals while expeditioning with [[Louis Leakey]] in the vicinity of [[Lake Victoria]]. The Consul that he selected to use in the name was neither of the ones mentioned above, but another located in the [[London Zoo]]. Consul is being used Linnaean-style to symbolize the [[Common chimpanzee|chimpanzee]]. ''Proconsul'' is therefore "ancestral to the Chimpanzee" in Hopwood's words. He also added ''africanus'' as the specific name.<ref name="Morell" />
 
Other fossils discovered later were initially classified as ''africanus'' and subsequently reclassified; that is, the total pool of fossils originally considered ''africanus'' was "split" and the fragments "lumped" with other finds to create a new species. For example, [[Mary Leakey]]'s famous find of 1948 began as ''africanus'' and was split from it to be lumped with Thomas Whitworth's finds of 1951 as ''heseloni'' by [[Alan Walker (anthropologist)|Alan Walker]] in 1993. This process creates some confusion for the public, which is told that ''africanus'' became ''heseloni''. The finds from Koru and Songhor are still considered ''africanus''. Four species are still defined even though many fossils have jumped species.<ref>{{Harvnb|Tuttle|2006| loc=''Taxonomic Shuffles, Ancestors, and Functional Interpretations: 1960–1999'', p. 17}}</ref>
 
The family of [[Proconsulidae]] was first proposed by Louis Leakey in 1963,<ref>{{Cite web | title =Proconsulidae | publisher = Palaeodatabase | url = http://paleodb.org/cgi-bin/bridge.pl?a=basicTaxonInfo&taxon_no=132667}}</ref> a decade after he and [[Wilfrid Le Gros Clark]] had defined ''africanus'', ''nyanzae'' and ''major''. It was not immediately accepted but ultimately prevailed.
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===Reassigned species===
The species ''Proconsul heseloni'' and ''P. nyanzae'' have been reclassified in the new genus ''[[Ekembo]]''.<ref>{{Cite journal|last1=McNulty|first1=Kieran P.|last2=Begun|first2=David R.|last3=Kelley|first3=Jay|last4=Manthi|first4=Fredrick K.|last5=Mbua|first5=Emma N.|date=2015|title=A systematic revision of Proconsul with the description of a new genus of early Miocene hominoid|journal=Journal of Human Evolution|language=en|volume=84|pages=42–61|doi=10.1016/j.jhevol.2015.03.009|pmid=25962549|doi-access=free|hdl=2286/R.I.35702|hdl-access=free}}</ref>
 
==Notes==
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|last2=Shipman |first2=Pat
|title=The Ape in the Tree: An Intellectual & Natural History of Proconsul
|url=https://books.google.com/books?id=DauBuaRsJl0C&lpgpg=PA3
|publisher=The Belknap Press of Harvard University Press| location=Cambridge, Massachusetts, London, England
|year=2005 |isbn=978-0-674-01675-0
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{{Taxonbar|from=Q131791}}
 
[[Category:Proconsulidae]]
[[Category:Prehistoric apes]] <!-- for when it is considered an ape -->
[[Category:Miocene primates of Africa]]
[[Category:Fossil taxa described in 1933]]